Ronald A. Markle Professor of Biology, Director, Pre-Medicine and Science Majors Breadcrumb About ram29 Ronald A. Markle Professor of Biology, Director, Pre-Medicine and Science Majors Department Affiliations Department of Biology External Affiliations Huck Institutes of the Life Sciences Address 236 East Ritenour Building, University Park, PA 16802 Email ram29@psu.edu Phone 814-865-7620
