**Tom will be arriving at Penn State in September 2022**
Ph.D., The University of Chicago, USA, 2015
B.S. Syracuse University, USA, 2009
Postdoctoral Scholar, The University of Chicago, USA 2017-2021
Postdoctoral Fellow, Yale University, USA, 2015-2017
Postdoctoral Fellow, University of Minnesota, USA, 2015-2017
I employ evolutionary and developmental approaches to ask: how does morphological novelty evolve, and what are the causes of major evolutionary transitions?
This work focuses on vertebrate appendages—fins and limbs—which are powerful models for testing hypotheses of developmental evolution and the relationship between form and function. By integrating paleontological data with experimental embryological and developmental genetic data, my aim is to discover how new body parts evolve and the biological basis of homology.